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Very similar.
The main differences are that those projects are highly configurable and can do a lot of things, while towonel is simpler: opinionated/streamlined for use as a shared Cloudflare tunnel alternative. I also think towonel may be the only one to use QUIC for the tunnel, just like Cloudflare.
Besides that, towonel is very new and still in alpha. Rathole does not seem to be actively developed anymore, which can be a good or bad thing.